B.A. Honors Thesis, Harvard University, Department of Biochemical Sciences: The Effects of Coprostanol and Other Related Sterols on the Growth and Membranes of Mycoplasma capricolum, Thesis Advisor: Professor Konrad Bloch, Ph.D., Harvard University (1981).

M.D. Honors Thesis, Harvard Medical School, Division of Health Sciences and Technology: Drug and Gene Delivery into the Central Nervous System, Thesis Advisor: Robert Langer,Sc.D., Massachusetts Institute of Technology (1990).

Ph.D. Thesis, Massachusetts Institute of Technology, Division of Health Sciences and Technology: Excitotoxic Mechanisms in Huntington’s Disease, Thesis Advisors: Joseph B. Martin, M.D., Ph.D. and Marian DiFiglia, Ph.D., Harvard Medical School and Massachusetts Institute of Technology (1992).

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